Llangefni is located just off the A55 Expressway in the centre of the island. It is the county town and administrative centre of Anglesey, and the County Council has its main office and council chamber there. It is an historic market town, with a stone clock tower in its central square, a feature shared by a number of other towns and villages on the island, such as Rhosneigr and Newborough.

Llangefni is home to Oriel Ynys Môn, the museum and arts centre, which was opened by Queen Elizabeth in October 1991. The Oriel's main gallery hosts a number of major exhibitions every year, many with a local focus.

One of the biggest attractions is the Sir Kyffin Williams Gallery, which opened in 2008. The County Council has over 400 of Sir Kyffin's works of art, and a number of these are always on display, together with other original works from other collections.

The Oriel is also home to a substantial collection of drawings and paintings by the famous wildlife artist Charles Tunnicliffe.

Key features of the town are the Town Hall or Neuadd y Dre, which was constructed in 1884 with limestone quarried on the eastern coast of Anglesey. It overlooks the central square or Maes Bulkley, with its stone tower and clock, also built from limestone.

At one side of the square is the impressive Bull Hotel or Gwesty'r Bull. A street market takes over the square and the adjoining car parks two days a week, Thursday and Saturday. Llangefni also has its own leisure centre at Plas Arthur.

On the outskirts of Llangefni is Nant y Pandy, or the Dingle. This is an extensive nature reserve comprising woodland and wetlands, with easy access over footpaths and boardwalks. One walk leads to Llyn Cefni, one of two reservoirs which supply drinking water to the people of Anglesey. This reservoir was created after the Second World War by damming the Afon Cefni. For the keen walker, it is possible to walk most of the way round the lake.

Instead of going around the reservoir, the railway took a more direct route of crossing it on an embankment and bridge.